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    John Greenland, MD, PhD

    TitleAssistant Professor
    SchoolUCSF School of Medicine
    Address4150 Clement St
    San Francisco CA 94121
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      Dr. Greenland is an Assistant Professor of Medicine at UCSF and Staff Physician at the San Francisco VA Medical Center. He received his undergraduate degree from Stanford and completed his PhD through the Harvard graduate program in Virology. He received an MD from the Harvard-MIT Division of Health Science and Technology.

      He serves as an attending physician in the intensive care unit, pulmonary consult service and outpatient pulmonary clinic at the SF VAMC. His research focuses on the immunology of lung transplant rejection and is primarily based in the analysis of human airway and peripheral blood samples. This research uses cellular immunology and genomic techniques to understand acute and chronic lung transplant rejection, with a goal of improving outcomes following lung transplantation.

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      Publications listed below are automatically derived from MEDLINE/PubMed and other sources, which might result in incorrect or missing publications. Researchers can login to make corrections and additions, or contact us for help.
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      1. Greenland JR, Wong CM, Ahuja R, Wang AS, Uchida C, Golden JA, Hays SR, Leard LE, Rajalingam R, Singer JP, Kukreja J, Wolters PJ, Caughey GH, Tang Q. Donor-Reactive Regulatory T-Cell Frequency Increases During Acute Cellular Rejection of Lung Allografts. Transplantation. 2016 Apr 13. PMID: 27077597.
        View in: PubMed
      2. Singer JP, Diamond JM, Gries CJ, McDonnough J, Blanc PD, Shah R, Dean MY, Hersh B, Wolters PJ, Tokman S, Arcasoy SM, Ramphal K, Greenland JR, Smith N, Heffernan P, Shah L, Shrestha P, Golden JA, Blumenthal NP, Huang D, Sonett J, Hays S, Oyster M, Katz PP, Robbins H, Brown M, Leard LE, Kukreja J, Bacchetta M, Bush E, D'Ovidio F, Rushefski M, Raza K, Christie JD, Lederer DJ. Frailty Phenotypes, Disability, and Outcomes in Adult Candidates for Lung Transplantation. Am J Respir Crit Care Med. 2015 Dec 1; 192(11):1325-34. PMID: 26258797.
        View in: PubMed
      3. Kern RM, Singer JP, Koth L, Mooney J, Golden J, Hays S, Greenland J, Wolters P, Ghio E, Jones KD, Leard L, Kukreja J, Blanc PD. Lung transplantation for hypersensitivity pneumonitis. Chest. 2015 Jun; 147(6):1558-65. PMID: 25412059; PMCID: PMC4451710 [Available on 06/01/16].
      4. Greenland JR, Xu X, Sayah DM, Liu FC, Jones KD, Looney MR, Caughey GH. Mast cells in a murine lung ischemia-reperfusion model of primary graft dysfunction. Respir Res. 2014; 15:95. PMID: 25115556; PMCID: PMC4151192.
      5. Greenland JR, Jewell NP, Gottschall M, Trivedi NN, Kukreja J, Hays SR, Singer JP, Golden JA, Caughey GH. Bronchoalveolar lavage cell immunophenotyping facilitates diagnosis of lung allograft rejection. Am J Transplant. 2014 Apr; 14(4):831-40. PMID: 24512389; PMCID: PMC4300200.
      6. Xu X, Greenland J, Baluk P, Adams A, Bose O, McDonald DM, Caughey GH. Cathepsin L protects mice from mycoplasmal infection and is essential for airway lymphangiogenesis. Am J Respir Cell Mol Biol. 2013 Sep; 49(3):437-44. PMID: 23600672; PMCID: PMC3824055.
      7. Greenland JR, Jones KD, Hays SR, Golden JA, Urisman A, Jewell NP, Caughey GH, Trivedi NN. Association of large-airway lymphocytic bronchitis with bronchiolitis obliterans syndrome. Am J Respir Crit Care Med. 2013 Feb 15; 187(4):417-23. PMID: 23239157; PMCID: PMC3603592.
      8. Geiben-Lynn R, Greenland JR, Frimpong-Boateng K, Letvin NL. Non-classical natural killer T cells modulate plasmid DNA vaccine antigen expression and vaccine-elicited immune responses by MCP-1 secretion after interaction with a beta2-microglobulin-independent CD1d. J Biol Chem. 2009 Dec 4; 284(49):33800-6. PMID: 19833737; PMCID: PMC2797149.
      9. Geiben-Lynn R, Greenland JR, Frimpong-Boateng K, van Rooijen N, Hovav AH, Letvin NL. CD4+ T lymphocytes mediate in vivo clearance of plasmid DNA vaccine antigen expression and potentiate CD8+ T-cell immune responses. Blood. 2008 Dec 1; 112(12):4585-90. PMID: 18784371; PMCID: PMC2597129.
      10. Geiben-Lynn R, Greenland JR, Frimpong-Boateng K, Letvin NL. Kinetics of recombinant adenovirus type 5, vaccinia virus, modified vaccinia ankara virus, and DNA antigen expression in vivo and the induction of memory T-lymphocyte responses. Clin Vaccine Immunol. 2008 Apr; 15(4):691-6. PMID: 18272665; PMCID: PMC2292663.
      11. Qin L, Greenland JR, Moriya C, Cayabyab MJ, Letvin NL. Effects of type I interferons on the adjuvant properties of plasmid granulocyte-macrophage colony-stimulating factor in vivo. J Virol. 2007 Oct; 81(19):10606-13. PMID: 17652387; PMCID: PMC2045443.
      12. Greenland JR, Geiben R, Ghosh S, Pastor WA, Letvin NL. Plasmid DNA vaccine-elicited cellular immune responses limit in vivo vaccine antigen expression through Fas-mediated apoptosis. J Immunol. 2007 May 1; 178(9):5652-8. PMID: 17442948; PMCID: PMC2262927.
      13. Greenland JR, Letvin NL. Chemical adjuvants for plasmid DNA vaccines. Vaccine. 2007 May 10; 25(19):3731-41. PMID: 17350735.
        View in: PubMed
      14. Hovav AH, Cayabyab MJ, Panas MW, Santra S, Greenland J, Geiben R, Haynes BF, Jacobs WR, Letvin NL. Rapid memory CD8+ T-lymphocyte induction through priming with recombinant Mycobacterium smegmatis. J Virol. 2007 Jan; 81(1):74-83. PMID: 17050608; PMCID: PMC1797252.
      15. Greenland JR, Liu H, Berry D, Anderson DG, Kim WK, Irvine DJ, Langer R, Letvin NL. Beta-amino ester polymers facilitate in vivo DNA transfection and adjuvant plasmid DNA immunization. Mol Ther. 2005 Jul; 12(1):164-70. PMID: 15963932.
        View in: PubMed
      16. Dominski Z, Erkmann JA, Greenland JA, Marzluff WF. Mutations in the RNA binding domain of stem-loop binding protein define separable requirements for RNA binding and for histone pre-mRNA processing. Mol Cell Biol. 2001 Mar; 21(6):2008-17. PMID: 11238936; PMCID: PMC86798.
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